Houbara Publications



Islam, M. Z., Boug, A., Alshariri, S., Alsubai, H. 2014. Houbara Bustard movement between Saudi Arabia and Kazakhstan. Phoenix, 30 (2014): 4-5.


Korrida, A. & Shweizer, M. 2013. Diversification across the Palaearctic desert belt throughout the Pleistocene: phylogeographic history of the Houbara - Macqueen’s bustrad complex (Otididae: Chlamydotis) as revealed by mitochondrial DNA. Journal of Zoological Systematics and Evolutionary Research 52 (1): 65-74.

Shobrak, M. Y., Hassan S. A. ., Stiévenart C., El-Deeb B. A. & Gherbawy Y. A. 2013. Prevalence and Antibiotic Resistance Profile of Intestinal Bacteria Isolated from Captive Adult Houbara Bustard (Chlamydotis macqueenii) Exposed to Natural Weather Conditions in Saudi Arabia. Global Veterinaria 10 (3): 276-284.

Korrida, A., Jadallah, S., Chbel, F., Amin-Alami, A., Ahra, M. & Aggrey, S. 2012. Patterns of genetic diversity and population structure of the threatened Houbara and Macqueen’s bustards as revealed by microsatellite markers. Genetics and Molecular Research 11 (3): 3207-3221.

Islam, M. Z., Singh A., Basheer M. P., Judas J. & Boug A. 2012.  Differences in space use and habitat selection between captive-bred and wild=born houbara bustards in Saudi Arabia: results from a long-term reintroduction program. Journal of Zoology, ZSL 2012.

Arif, I.A., Khan, H. A., Williams, J. B., Shobrak, M. & Arif, W. I. 2012. DNA Barcodes of Houbara Bustard (Chlamydotis undulata macqueenii). International Journal of Molecular Sciences, 2012, 13: 2425-2438.

Islam, M.Z. Basheer P.M., Rahman W. and Boug, A. 2011. An attack by Ratel Mellivora capensis on pre-release Asian Houbara Bustards Chlamydotis macqueenii in central Saudi Arabia. Small Carnivore Conservation, June 2011, Vol. 44: 35-37.

Islam, M.Z., Boug, A. 2011. Save the Houbara... Hornbill, January-March 2011: 18-21.

Islam, M. Z., Boug, A., Basheer, M. S., Shah, M. S., & Subai, H. 2010. Reintroduction successes of Asian Houbara Bustard Chlamydotis macqueenii in the Kingdom of Saudi Arabia. IUCN: Global Reintroduction Perspectives: 132-138.

Bailey, TA., McKinney, P. & Stiévenart, C. 2008. Neonatal Diseases. In Disease and medical management of houbara bustards and other Otididae. Bailey TA Ed. (Emirates Printing Press LLC, Dubai): 301-317.


Stiévenart, C. 2008. A novel, non-surgical method for the treatment of tibio-tarsal rotation in Asian houbara bustard Chlamydotis macqueenii (Otididae) chicks. Journal of Avian Medicine and Surgery. 22(1): 25-30.

Islam, M. Z., Basheer P.M., Shah M.S.; Subai H., Boug, A. 2008. Conservation challenges to continue the reintroduction of Houbara bustard Chlamydotis macqueenii in the Kingdom of Saudi Arabia. FALCO The Newsletter of the Middle East Falcon Research Group (No. 32).


Stiévenart, C. 2007. Health problems during the first month of life of captive-born Asian Houbara bustard Chlamydotis macqueenii chicks.  Proceedings of the 9th European Association of Avian Veterinarian EAAV Conference (March 2007, Zurich): 105-113.


Islam, M. Z. 2007. Return of the native: re-introducing the houbara bustard to Saudi Arabia. Species - Newsletter of the Species Survival Commission - SSC(47): 3.


Islam, M. Z., Basheer, P.M., Shah, M.S., Al-Suabi, H. 2007. The houbara bustard in the Kingdom of Saudi Arabia. CBSG News, 18(2): 1.


Islam, M. Z., Shah, M.S., Basheer, P.M.,  Al-Suabi, H., Shobrak, M. 2007. Houbara bustard re-introduction in the Kingdom of Saudi Arabia. Re-introduction NEWS(26): 3.


Islam, M. Z., Basheer, P.M., Shah, M.S., Al-Suabi, H., Shobrak, M. 2007. Conservation efforts to restore the Houbara bustard Chlamydotis macqueenii population in the Kingdom of Saudi Arabia. Wildlife Middle East, 2(2): 2.


Hossain, J., Stiévenart, C., Hémon, S., Staines, H.J., & Wishart, G.J. 2005. Male-specific differences in the nuclear and flagellar lengths of spermatozoa from Asian houbara bustards Chlamydotis macqueenii. Avian and Poultry Biology Reviews, 16(1): 55-56.

Stiévenart, C. 2004. Laying pattern and egg mass of captive Asian houbara bustards Chlamydotis macqueenii subjected to egg-removal. Avian and Poultry Biology Reviews, 16(1): 69.


Stiévenart, C. & Mohammed, OB. 2004. Current and emerging diseases in a captive flock of adult houbara bustards (Chlamydotis macqueenii). Proceedings of the 3rd Annual Meeting of the Africa and Middle East Section of the Wildlife Diseases Association WDA (Abu Dhabi, December 2004).


Maloney, R. F. 2004. [Asian houbara bustard / ecology / Mahazat as-Sayd]. Unpublished PhD Thesis.


Broders O., Osborne T. & Wink M. 2003. A mtDNA phylogeny of bustards (family Otididae) based on nucleotide sequences of the cytochrome b-gene. J. Ornithol. 144: 176-185.


Stiévenart, C. 2002. Monitoring of growth in houbara bustard Chlamydotis [undulata] macqueenii chicks.  Proceedings of the 27th World Veterinary Congress (September 2002, Tunis): 178-179.

van Heezik, Y., & Seddon, P. J. 2002. Patch use and exploratory movements of a resident houbara bustard in northern Saudi Arabia. Journal of Arid Environments, 50(4): 683-686.


van Heezik, Y., Saint Jalme, M., Hémon, S., & Seddon, P. J. 2002. Temperature and egg-laying experience influence breeding performance of captive female houbara bustards. Journal of Avian Biology, 33(1): 63-70.


Tieleman, B. I., Williams, J. B., Lacroix, F., & Paillat, P. 2002. Physiological responses of houbara bustards to high ambient temperatures. The Journal of Experimental Biology, 205(4): 503-511.


Tieleman, B. I. 2002. Physiological responses of Houbara bustards to high ambient temperatures. Irene Tieleman (ed.) Avian Adaptation Along an Aridity Gradient. PhD Groningen University: 247.


Judas, J. 2002. Réintroduction de l'outarde houbara Chlamydotis [undulata] macqueenii en Arabie Saoudite. Alauda. Actes du 26e Colloque National Francophone d'Ornithologie, 70(1): 180-181.


Judas, J. & Routier, J-B. 2002. Survival parameters of a reintroduced Asian Houbara Bustard population in central Saudi Arabia. International Ornithological Congress, Beijing, China.


Baggot, G. K., Deeming, D. C., Hémon, S., & Paillat, P. 2002. Relationships between eggshell pigmentation, ultrastructure and water vapour conductance in the houbara bustard (Chlamydotis undulata macqueenii). Avian and Poultry Biology Reviews. WPSA Working Group 6 (Reproduction). Abstracts of the Incubation and Fertility Research Group Meeting., 13(4): 234-235.


van Heezik, Y., & Seddon, P. J. 2001. Influence of group size and neonatal handling on growth rates, survival, and tameness of juvenile houbara bustards. Zoo Biology, 20(5): 423-433.


van Heezik, Y., & Seddon, P. 2001. Born to be tame. Natural History, 6: 58-63.


van Heezik, Y., & Ostrowski, S. 2001. Conservation breeding for reintroductions: assessing survival in a captive flock of houbara bustards. Animal Conservation, 4(3): 195-201.


Maloney, R. F. 2001. Sexing of houbara bustards Chlamydotis [undulata] macqueenii, using footprint measurements. Ostrich, 72(3-4): 216-218.


Lenain, D. M., & Warrington, S. 2001. Is translocation an effective tool to remove predatory foxes from a desert protected area? Journal of Arid Environments, 48(2), 205-209.


Judas, J., & Paillat, P. 2001. Re-introduction program of houbara bustard in Saudi Arabia. Houbara News, 4: 4.


Deeming, D. C., Paillat, P., Hémon, S., & Saint Jalme, M. 2001. Attentiveness and turning patterns during incubation in a houbara bustard (Chlamydotis undulata macqueenii) nest. Avian and Poultry Biology Reviews. WPSA Working Group 6 (Reproduction). Abstracts of the Incubation and Fertility Research Group Meeting., 12(4): 182-183.


Soorae, P. S., & Seddon, P. J. 2000. (article on houbara reintroduction). Re-introduction News. Special Bird Issue. Newsletter of the IUCN/SSC Re-introduction Specialist Group, 20, 56 pp.


Seddon, P. J., Judas, J., Maloney, R., Combreau, O., & van Heezik, Y. 2000. The re-introduction of houbara bustards in the Kingdom of Saudi Arabia. Re-introduction News. Special Bird Issue. Newsletter of the IUCN/SSC Re-introduction Specialist Group, 19: 22-24.


Seddon, P. J. 2000. The re-introduction of houbara bustards in the Kingdom of Saudi Arabia. Re-Introduction News: 22-24.


Judas, J. 2000. Reintroduction of houbara bustard in central Saudi Arabia. British Poultry Science. Special Issue: Proceedings of the International Conference on Bird Reproduction, Tours, France, September 1999. Volume 41, Supplement 1, S50.


Judas, J. 2000. L'outarde houbara. Le Courrier de la Nature, 182: 44-45.


Hémon, S., Paillat, P., van Heezik, Y., & Judas, J. 2000. Captive breeding of houbara bustard in Saudi Arabia: 11 successful years. Bristish Poultry Science. Special Issue: Proceedings of the International Conference on Bird Reproduction, Tours, France, September 1999., Volume 41, Supplement 1, S49-S50.


Combreau, O., Gélinaud, G., & Smith, T. R. 2000. Home range and movements of houbara bustards introduced in the Najd pediplain in Saudi Arabia. Journal of Arid Environments, 44(2): 229-240.


van Heezik, Y., Seddon, P. J., & Maloney, R. F. 1999. Helping reintroduced houbara bustards avoid predation: effective anti-predator training and the predictive value of pre-release behaviour. Animal Conservation, 2(3): 155-163.


van Heezik, Y., & Seddon, P. J. 1999. Seasonal changes in habitat use by houbara bustards Chlamydotis [undulata] macqueenii in northern Saudi Arabia. Ibis, 141: 208-215.


Seddon, P. & Van Heezik, Y. 1999. Counting Houbara Bustard in northern Saudi Arabia: An assessment of method. 18-24. (Launay, F. and Bailey, T., Eds). Counting Houbara Bustard. NARC, IUCN/SSC/BirdLife International, Cambridge.


Seddon, P. J., Launay, F., van Heezik, Y., & Al Bowardi, M. 1999. Methods for live trapping houbara bustards. Journal of Field Ornithology, 70(2): 169-181.


Seddon, P. J. 1999. Persistence without intervention: assessing success in wildlife reintroductions. Trends in Ecology and Evolution, 14(12): 503.

Seddon, P. J. 1999. Arabia's bustard rescued from falconers. Endangered species, 1(2): 65-67.


Judas, J. 1999. Reintroduction of houbara bustard in central Saudi Arabia. Science, B. P. (ed.) International Conference on Bird Reproduction: 50.


van Heezik, Y., Seddon, P. J., & Maloney, R. F. 1998. Behavioural preparation of captive-bred houbara bustards for re-introduction. Proceedings of the 22nd International Ornithological Congress, Durban, South Africa. Ostrich, 69: 270.


van Heezik, Y., & Seddon, P. J. 1998. Ontogeny of behaviour of hand-reared and hen-reared captive houbara bustards. Zoo Biology, 17(3): 245-255.


Seddon, P. J. 1998. Improving the rigour of reintroduction project assessment, planning and execution: report on the round table discussions on bird re-introductions. Re-introduction News. Newsletter of the IUCN/SSC Re-introduction Specialist Group, 16: 3-5.

Ostrowski, S., Saint Jalme, M., & Ancrenaz, M. 1998. Antibody response to Newcastle disease vaccination in a flock of young houbara bustard (Chlamydotis undulata). Journal of Zoo and Wildlife Medicine, 29(2): 234-236.


Maloney, R. F., Seddon, P. J., & Paillat, P. 1998. Reintroduction of houbara bustards: the process of establishing a resident breeding population. Proceedings of the 22nd International Ornithological Congress, Durban, South Africa. Ostrich, 69: 239.


Maloney, R. F. 1998. Video monitoring observations of a houbara bustard Chlamydotis undulata nest in Mahazat as-Sayd reserve, central Saudi Arabia. Sandgrouse, 20(1): 36-39.


Hémon, S. 1998. Raising the houbara bustard. Game Bird and Conservationists' Gazette, October 1998: 39-40.


Hémon, S. 1998. Captive propagation of houbara bustards in Saudi Arabia: 1998 breeding season. WAWV Newsletter, 7(32): 3-4.


Hémon, S. 1998. Houbara bustard captive breeding update: more than 240 chicks produced and 100 to be released in Saudi Arabia. Re-introduction News. Newsletter of the IUCN/SSC Re-introduction Specialist Group: 16, 6.


Hémon, S. 1998. Houbara captive breeding program in Taif, Saudi Arabia. WAWV Newsletter, 7(30): 5.


Combreau, O., & Smith, T. R. 1998. Release techniques and predation in the reintroduction of houbara bustards in Saudi Arabia. Biological Conservation, 84: 147-155.


Cade, T. J., & Seddon, P. J. 1998. Bird re-introductions: pre-release preparations, releases and post-release monitoring. Proceedings of the 22nd International Ornithological Congress, Durban, South Africa. Ostrich, 69: 161.


van Heezik, Y., & Maloney, R. F. 1997. Update on the houbara bustard reintroduction programme in Saudi Arabia. Re-introduction News. Newsletter of the IUCN/SSC Re-introduction Specialist Group, 13: 3-4.


Seddon, P. J., & Abuzinada, A. H. 1997. Saudi Arabia expands its network of protected areas for migratory houbara bustards. Houbara News, 2: 4-6.


Seddon, P. J. 1997. Resident houbara bustard populations in Saudi Arabia: do summer ambient temperatures limit distribution? Journal of Arid Environments, 37(3): 551-556.


Pierrard, P. 1997. The Taif National Wildlife Research Center or the struggle against final emptiness in the desert. Risalat al-Ma'ahad, 10(104-105): 26-29.


Ostrowski, S. 1997. L'outarde houbara et la variole aviaire [The houbara bustard and the avian pox disease]. Unpublished Thèse de Doctorat Vétérinaire [ Veterinary Thesis], Ecole Nationale Vétérinaire de Lyon, Marcy l'Etoile: 155pp.

Gélinaud, G., Combreau, O., & Seddon, P. J. 1997. First breeding success in captive-born reintroduced houbara bustards. Journal of Arid Environments, 35(3): 527-534.


Combreau, O., & Smith, T. R. 1997. Summer habitat selection by houbara bustards introduced in central Saudi Arabia. Journal of Arid Environments, 36(1): 149-160.


Besson, F. 1997. Le Centre National de Recherche sur la Vie Sauvage de Taif. Nouvelles Scientifiques de France et du Proche-Orient, Mars, 80-83.


Bailey, F., & Launay, F. 1997. Satellite tracking houbara bustards. Arabian Wildlife, 3(2): 14-16.


Anon. 1997. Houbara bustard are satellite tracked, from the UAE to Asia. The Phoenix, 14: 17-18.


Ady, J. 1997. The National Wildlife Research Center, at Ta'if. Journal of the Saudi Arabian Natural History Society, 3(7): 1-3.


van Heezik, Y., & Seddon, P. J. (Eds.). 1996. Restoration of bustard populations: captive breeding, release, monitoring and habitat management. Riyadh: NCWCD Publication No.27, 111pp.

Seddon, P. J., & van Heezik, Y. 1996. Seasonal changes in houbara bustard Chlamydotis undulata macqueenii numbers in Harrat al Harrah, Saudi Arabia: implications for managing a remnant population. Biological Conservation, 75(2): 139-146.


Seddon, P. J., & Maloney, R. F. 1996. Reintroduction of houbara bustards into central Saudi Arabia: a summary of results between 1991 and 1996. The Phoenix, 13: 14-16.


Saint Jalme, M., & van Heezik, Y. (Eds.). 1996. Propagation of the houbara bustard. London/Riyadh: Kegan Paul International/NCWCD, 110pp.

Saint Jalme, M., Williams, J., Mikaelian, I., Gaucher, P., & Paillat, P. 1996. Seasonal variation of LH, sex steroids, body mass, moult, display and laying in two subspecies of houbara bustard Chlamydotis undulata undulata and Chlamydotis undulata macqueenii housed in outdoor cages under natural conditions. General and Comparative Endocrinology, 102(1): 102-112.


Saint Jalme, M., Paillat, P., Hemons, S. & Ostrowski, S. 1996. Importance of Studies on Breeding Biology in the Conservation Program of Houbara Bustard in Saudi Arabia. Gutiérrez, J. F. and Sanz-Zuasti, J. (eds.), Conservation de las Aves Esteparias y su Habitat. Juan de Castilla y Leon: 161-170.


Saint Jalme, M., Combreau, O., Seddon, P. J., Paillat, P., Gaucher, P., & van Heezik, Y. 1996. Restoration of Chlamydotis undulata macqueenii (houbara bustard) populations in Saudi Arabia: a progress report. Restoration Ecology, 4(1): 1-8.


Ostrowski, S., Dorrestein, G., Burger, L., Hémon, S., & Saint Jalme, M. 1996. Cross-protection test of an avian poxvirus isolated from houbara bustards. Avian Diseases, 40(3): 762-769.


Gubin, B. M. 1993, 1996. Houbara bustards in Kazakhstan: distribution, numbers, seasonal movements, research effort and protection. Paper presented at the Restoration of Bustard Populations. Proceedings of the Symposium Restoration of Houbara Bustard Populations in Saudi Arabia, Taif and Riyadh (29 September-4 October 1993, 1996): 15-19.

Gaucher, P., Paillat, P., Chappuis, P., Saint Jalme, M., Lotfikhah, F., & Wink, M. 1996. Taxonomy of the houbara bustard Chlamydotis undulata subspecies considered on the basis of sexual display and genetic divergence. Ibis, 138(2): 273-282.


Combreau, O., & Launay, F. 1996. Activity rhythms of houbara bustards (Chlamydotis undulata macqueenii) in relation to some abiotic factors. Journal of Arid Environments, 33(4): 463-472.


Al-Saihati, A. W. 1996. Studies and conservation of houbara bustard (Chlamydotis undulata macqueenii) in Saudi Arabia. Unpublished Doctor of Philosophy Thesis, The Union Institute, Cincinnati, 327pp.

Seddon, P. J., Saint Jalme, M., van Heezik, Y., Paillat, P., Gaucher, P., & Combreau, O. 1995. Restoration of houbara bustard populations in Saudi Arabia: developments and future directions. Oryx, 29(2): 136-142.


Seddon, P. J., & Gélinaud, G. 1995. Reintroduced houbara bustards begin to breed in the Mahazat as-Sayd Reserve, Saudi Arabia. Re-introduction News. Newsletter of the IUCN/SSC Re-introduction Specialist Group, 11: 6-7.


Seddon, P. J. 1995. Born to be wild: reintroduced houbara bustards breed for the first time in Saudi Arabia. Arabian Wildlife, 2(3): 16-18.


Ostrowski, S., Dorrestein, G., Ancrenaz, M., & Saint Jalme, M. 1995. Debilitating cutaneous poxvirus lesions on two captive houbara bustards (Chlamydotis undulata). Avian Diseases, 39(4): 907-911.


Ostrowski, S. and Combreau, O. 1995. A survey of causes of mortality in captive, captive bred released, and wild born houbara bustards (Chlamydotis undulata) in Saudi Arabia. - In: 87-97.


Ostrowski, S., Ancrenaz, M., Saint Jalme, M., & Greth, A. 1995. Concurrent avian pox and Newcastle disease infection in a houbara bustard (Chlamydotis undulata). Avian Pathology, 24: 573-577.


Nader, I. A. 1995. Harrat al-Harrah. First national reserve in Saudi Arabia. Arabian Wildlife, 2(2): 14.


Gaucher, P. 1995. Breeding biology of the houbara bustard Chlamydotis undulata in Algeria. Alauda, 63(4): 291-298.


Combreau, O., Saint Jalme, M., Seddon, P. J., Rambaud, F., van Heezik, Y., Paillat, P., Gaucher, P. & Smith, T. 1995. A Program for Houbara Bustard Restoration in Saudi Arabia. Bissonet, J. A. and Krausman, P. R. (eds.), Integrating people and wildlife for a sustainable future. Proceeding of the first International Wildlife Management Congress. The Wildlife Society: 697.


Saint Jalme, M., Gaucher, P., & Paillat, P. 1994. Artificial insemination in houbara bustards (Chlamydotis undulata): influence of the number of spermatozoa and insemination frequency on fertility and ability to hatch. Journal of Reproduction and Fertility, 100: 93-103.


Saint Jalme, M. 1994. Houbara: the Saudi Arabian project. Arabian Wildlife, 1(2), 6-8.


Liedeker, H., Al Welaie, A. N., & As Saifi, A. 1994. GIS Can they deliver for conservation? Arabian Wildlife, 1(2): 12-13.


Granjon, L., Gaucher, P., Greth, A., Paillat, P., & Vassart, M. 1994. Allozyme study of two subspecies of houbara bustards (Chlamydotis undulata undulata). Biochemical Systematics and Ecology, 22(8): 775-779.


Anon. 1994. Update: Reintroduction of houbara bustard in Saudi Arabia. Arabian Wildlife, 1(2): 39.


Anegay, K. 1994. Diurnal and nocturnal activity patterns of semi-captive houbara bustards Chlamydotis undulata. Sandgrouse, 16(1): 41-46.


Abuzinada, A. H., & Seddon, P. J. 1994. Report on the NCWCD workshop on the restoration of houbara bustard populations in Saudi Arabia. Re-introduction News. Newsletter of the IUCN/SSC Re-introduction Specialist Group, 8: 2-3.


Mikaelian, I. 1993. Variations circannuelles des paramètres hématologiques de l'outarde houbara (Chlamydotis undulata). Unpublished Thèse de Doctorat Vétérinaire [ Veterinary Thesis], Ecole Nationale Vétérinaire de Lyon, Marcy l'Etoile, 149pp.

Greth, A., Gerlach, H., Gerbermann, H., Vassart, M., & Richez, P. 1993. Pharmacokinetics of doxycycline after parenteral administration in the houbara bustard Chlamydotis undulata. Avian Diseases, 37(1): 31-36.


Greth, A., Andral, B., Gerbermann, H., Vassart, M., Gerlach, H., & Launay, F. 1993. Chlamydiosis in a captive breeding group of houbara bustards (Chlamydotis undulata). Avian Diseases, 37(4): 1117-1120.


Schulz, M., & Schulz, H. (Eds.). 1991. Working bibliography of the bustards (Otididae). Riyadh: NCWCD, 171pp.

Gaucher, P. 1991. On the feeding ecology of the houbara Chlamydotis undulata undulata. Alauda, 59(2): 120-121.


Symens, P. 1990. Effects of the mass migration of desert locusts Schistocerca gregaria on birds in the Taif area, Saudi Arabia. Sandgrouse, 12: 3-7.


Launay, F., & Paillat, P. 1990. A behavioural repertoire of the adult houbara bustard (Chlamydotis undulata macqueenii). Revue d 'Ecologie - La Terre et la Vie, 45(1): 65-88.


Stephenson, P. 1988. The National Wildlife Research Center, Taif. Journal of the Saudi Arabian Natural History Society, 2(8), 54-58.


Jennings, M. C. 1988. The former distribution of the houbara bustard. The Phoenix, 5: 4.


Asmodé, J.-F. 1986. Taif. Centre de Recherche sur la Faune Sauvage [Taif Wildlife Research Center]. Le Courrier de la Nature, 106, 20-26.